Plant Cells


Plant Cells and Transpiration

Cells are the basic unit of plant life. Plant cells generally combine into larger subgroups called tissues. There are two major types of plant tissues: xylem and phloem. Xylem consists mainly of dead cells and serves to transport water. Phloem consists of live cells and transport food substances. These two types of tissues combine into organs. There are three main organs in the plant: stem, root and leaves. The focus of this article is on the xylem tissue. Xylem cells run continuously from the leaves to the roots. Their role is to conduct water from roots to the leaves. In this article, we shall review the mechanism by which plants take up water from the soil through its roots.

Contrary to what may be expected, the mechanism of water uptake in plants begins not at the roots, but at the leaves. In contact with the dry atmosphere, moisture from the leaves evaporate via the tiny pores located on the leaf surface. The loss of water creates an osmotic gradient. Osmosis is the process by which water moves from an area where it is in abundance to an area where it is scarce. Thus, since an osmotic gradient is created from the leaf tissues to the atmosphere, water vapor will leave the cells of the leaf surface for the atmosphere. Since the surface cells experience a water shortage, an osmotic gradient is again established between the water-poor surface cells and the water-rich deeper tissues. A chain of osmotic gradients is thus created from cell to cell causing water to move from the deeper layers to the surface. This osmotic pull is ultimately felt by the xylem cells, causing water to move up the xylem tissue into the leaves.

The osmotic gradient finally reaches the root cells and the procedure is repeated until the surface cells of the root are drained of water as well. By osmosis therefore, water surrounding the roots moves into the root cells. Through this mechanism water is taken up from the roots into the entire plant organism. This entire process is called transpiration.

It must be noted that transpiration is entirely passive. Energy is not used at all. Furthermore, through this process, the plant absorbs only the quantity of water that its cells need. If its cells are turgid (i.e. contain enough water), there will be no osmotic gradient and no water uptake.

If the atmosphere is saturated with water (100% humidity levels), then there will be no osmotic gradient created as well. Transpiration will not occur in this case. The higher the humidity level, the lower is the rate of transpiration. Conversely, a low humidity level increases the water uptake as the osmotic gradient is very steep. Under such conditions, if the soil contains insufficient water, transpiration will result in an overall loss of water. This results in wilting of the plant and may lead to death if such conditions persist long enough. Some plants, especially desert plants such as the cactus, have adapted themselves to prevent such unbridled transpiration. Their leaf pores close down under intense heat.

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